To serve the 397B-parameter Qwen 3.5 Mixture-of-Experts (MoE) model on Ironwood TPUs, engineers developed a modular JAX/Pallas optimization stack that achieved up to a 4.7x inference speedup for prefill-heavy workloads. The team bypassed severe hardware sharding constraints by deploying a hybrid Data Parallelism and Expert Parallelism (DP+EP) topology, paired with custom low-level communication fusions like a hierarchical reduce-scatter to optimize cross-device token routing. Finally, by executing hardware-aware custom kernels—such as Batched Ragged Page Attention and a fully-fused Gated DeltaNet (GDN) block—they successfully saturated HBM bandwidth and TensorCore MXUs to push system throughput near its theoretical roofline limits.